Mamadu Bama was the second-in-command, the military said in a statement.
He was killed this month along with his father, who was the group's spiritual mentor, the military said.
The militant group has not released any statement on the deaths.
Boko Haram has attacked
various targets in the nation since 2009, murdering and kidnapping
Westerners, and bombing schools and churches.
It is suspected to be
behind a gruesome mosque killing this week. In that attack Sunday,
militants brandishing automatic weapons killed 44 worshipers in the
country's troubled Borno state. Around the time of the attack, the group
released a video boasting that it was growing stronger.
Boko Haram translates
loosely to "western education is forbidden/sinful." It holds all
government authority in contempt and wants to establish a state in
northern Nigeria under Islamic law.
Human rights groups have accused the Nigerian military of extrajudicial killings in its fight against the militant group.
"Soldiers have allegedly
burned homes and executed Boko Haram suspects or residents with no
apparent links to the group," Human Rights Watch said in a report this year. "Nigerian authorities have rarely brought anyone to justice for these crimes."
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